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A wave breaks at sunset on the Brittany coast at Penmarch in Finistere
More than 20,000 seabirds died during the storms that hit France's west coast in January and February, according to the French society for the protection of birds. So many avain deaths have not been since 1900, the organisation says.
A series of storms hit the French coast along the Bay of Biscay and Brittany in January and February.
They caused extensive damage on land but also had a devastating effect on wildlife, the LPO reports, pointing out that the figure is probably an underestimate, given that sailors have reported seeing many birds' bodies floating at sea.
The worst-affected species were the Atlantic puffin and the guillemot with the razor-billed auk also suffering badly.
Most of the casualties were due to a lack of food because storms blew the birds to areas where less was available.
Others may have died of exhaustion because giant waves prevented them settling on the water and forced them to use more energy than usual, while the LPO suspects that some boats may have emptied oil under cover of the storm, further endangering seabirds.